Buildings groan as their innards expand and contract; joints between walls and rooflines separate. Where they do, rain leaks in, bolts rust, and facing pops off, exposing insulation. If the city hasn’t burned yet, it will now. Collectively, New York architecture isn’t as combustible as, say, San Francisco’s incendiary rows of clapboard Victorians. But with no firemen to answer the call, a dry lightning strike that ignites a decade of dead branches and leaves piling up in Central Park will spread flames through the streets. Within two decades, lightning rods have begun to rust and snap, and roof fires leap among buildings, entering paneled offices filled with paper fuel. Gas lines ignite with a rush of flames that blows out windows. Rain and snow blow in, and soon even poured concrete floors are freezing, thawing, and starting to buckle. Burnt insulation and charred wood add nutrients to Manhattan’s growing soil cap. Native Virginia creeper and poison ivy claw at walls covered with lichens, which thrive in the absence of air pollution. Red-tailed hawks and peregrine falcons nest in increasingly skeletal high-rise structures.

Within two centuries, estimates Brooklyn Botanical Garden vice president Steven Clemants, colonizing trees will have substantially replaced pioneer weeds. Gutters buried under tons of leaf litter provide new, fertile ground for native oaks and maples from city parks. Arriving black locust and autumn olive shrubs fix nitrogen, allowing sunflowers, bluestem, and white snakeroot to move in along with apple trees, their seeds expelled by proliferating birds.

Biodiversity will increase even more, predicts Cooper Union civil engineering chair Jameel Ahmad, as buildings tumble and smash into each other, and lime from crushed concrete raises soil pH, inviting in trees, such as buckthorn and birch, that need less-acidic environments. Ahmad, a hearty silver-haired man whose hands talk in descriptive circles, believes that process will begin faster than people might think. A native of Lahore, Pakistan, a city of ancient mosaic-encrusted mosques, he now teaches how to design and retrofit buildings to withstand terrorist attacks, and has accrued a keen understanding of structural weakness.